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Sinornithosaurus

Sunday, December 11, 2016

Sinornithosaurus millenii fossil displayed in Hong Kong Science Museum

Sinornithosaurus (derived from a combination of Latin and Greek, meaning ‘Chinese bird-lizard’) is a genus of feathered dromaeosaurid dinosaur from the early Cretaceous Period (early Aptian) of the Yixian Formation in what is now China. It was the fifth non–avian feathered dinosaur genus discovered by 1999. The original specimen was collected from the Sihetun locality of western Liaoning. It was found in the Jianshangou beds of the Yixian Formation, dated to 124.5 million years ago. Additional specimens have been found in the younger Dawangzhangzi bed, dating to around 122 million years ago.

Sinornithosaurus by PrehistoricWildlife.com

The small predatory dinosaur Sinornithosaurus was a dromaeosaurid-one of a group of agile bipedal runners with large eyes, relatively large brains, and long, narrow snouts equipped with steak-knife teeth. The three fingers on each hand and the four toes of each foot had long, curved, wickedly sharp claws for hooking into prey, with the second toe claw being extra large. As in many other dinosaurs, especially fast-running kinds, the tail was stiffened by overlapping bony rods.

Skeletal reconstructions of S. millenii and NGMC 91. Author: Jaime A. Headden

Sinornithosaurus was discovered by Xu Xing, Wang Xiaolin and Wu Xiaochun of the Institute of Vertebrate Paleontology and Paleoanthropology of Beijing. An almost-complete fossil with feather impressions, was recovered from Liaoning Province, China, in the Yixian Formation; the same incredibly rich location where four dinosaurs with feathers were discovered previously, ProtarchaeopteryxSinosauropteryxCaudipteryx, and Beipiaosaurus. The holotype specimen is IVPP V12811, in the collection of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, China.

The NGMC 91 specimen is in the collection of the National Geological Museum of China. It was collected in Fanzhangzi quarry, near Lingyuan City, Liaoning Province. This location is part of the Dawangzhangzi fossil beds, which have been dated to about 122 million years ago, during the early Aptian age. A specimen of the fish Lycoptera is also preserved near the foot.

Artist's impression of S. millenii

Phylogenetic studies did not support the idea that NGMC 91 was a close relative of S. millenii. In a 2004 analysis, Phil Senter and colleagues found that it was, in fact, more closely related to Microraptor. Subsequent studies, also by Senter, have continued to show support for this finding despite the fact that some data used in the original study was later found to be flawed.

Sinosauropteryx

Sunday, December 11, 2016

Sinosauropteryx

Sinosauropteryx (meaning “Chinese reptilian wing”, Chinese: 中华龙鸟; pinyin: Zhōnghuá lóng niǎo; literally: “China dragon bird”) is a compsognathid dinosaur. Described in 1996, it was the first dinosaur taxon outside of Avialae (birds and their immediate relatives) to be found with evidence of feathers. It was covered with a coat of very simple filament-like feathers. Structures that indicate colouration have also been preserved in some of its feathers, which makes Sinosauropteryx the first non-avialian dinosaurs where colouration has been determined. The colouration includes a reddish and light banded tail. Some contention has arisen with an alternative interpretation of the filamentous impression as remains of collagen fibres, but this has not been widely accepted.

Size of adult and sub-adult specimens, compared with a human. Author: Matt Martyniuk

Sinosauropteryx caused a sensation when it was revealed to the world in 1996. Here was a clear and perfectly preserved skeleton of a small theropod dinosaur, but covering most of the body were impressions of an enigmatic fuzz, a coat of fine filaments up to 1 1/2 inches (4 cm) long.

Skin impressions found for some dinosaurs reveal that they had scales. But the growing recognition that birds evolved from dinosaurs raised the possibility that some dinosaurs had feathers, or at least some kind of protofeather-like body covering that later evolved into the feathers of present-day birds. However, feathers are relatively soft and rarely fossilized.

Skeletal restoration of two specimens. Author: Jaime A. Headden

It is likely that several birdlike dinosaurs may have had feathers or similar structures, but, as they were not preserved, we know nothing about them. Then Sinosauropteryx was found in the fabulous Liaoning deposits in China. Here was the right kind of dinosaur (a small, advanced theropod) with much of its back, rump, and tail covered in a fine, filamentous fuzz that appeared to be fur-like coat made up of short, single strands.

While lacking the complex structure of a feather, it was a more intricate body covering than simple reptilian scales. The fuzzy coat seemed to form a downy layer that would have been perfect for trapping body heat and keeping the animal warm. This observation lends more weight to the ongoing debate that some dinosaurs were warm-blooded. It is also possible that the fuzz was used when displaying to attract mates.

Sinosauropteryx fossil, from our trip to Hohhot, Inner Mongolia, China for National Day Holiday 2007. This is the Inner Mongolia Museum, the biggest museum in the region. Sam / Olai Ose / Skjaervoy from Zhangjiagang, China

In one of the specimens was the remains of its last meal, a small, unidentified animal-providing proof that this little ancestor of the birds ate the ancestors of modern mammals. One of the other specimens revealed unlaid eggs in the oviducts.

Sinosauropteryx Photo: PA

Scutellosaurus

Sunday, December 11, 2016

Scutellosaurus

Scutellosaurus is an genus of thyreophoran ornithischian dinosaur that lived approximately 196 million years ago during the early part of the Jurassic Period in what is now Arizona, USA. It is classified in Thyreophora, the armoured dinosaurs; its closest relatives may have been Emausaurus and Scelidosaurus, another armored dinosaur which was mainly a quadrupedal dinosaur, unlike bipedal Scutellosaurus. It is one of the earliest representatives of the armored dinosaurs and the basalmost form discovered to date. Scutellosaurus was a small, lighly-built, ground-dwelling, herbivore, that could grow up to an estimated 1.2 m (3.9 ft) long.

1. Scelidosaurus harrisonii (“limb lizard”) 1861 Chordata/Ornithischia/Scelidosauridae. 2. Scutellosaurus lawleri (“little shielded lizard”) 1981 Chordata/Ornithischia.

The holotype specimen of Scutellosaurus lawleri (MNA V175) was recovered at the West Moenkopi Plateau locality in the Silty Facies Member of the Kayenta Formation, in Coconino County, Arizona on the land of the Navajo Nation. The specimen was discovered and collected by David Lawler in red claystone sediments that were deposited during the Sinemurian stage of the Jurassic period, approximately 196 million years ago.

Scutellosaurus is one of the earliest representatives of the armored dinosaurs (the thyreophorans) that would later include giants as Stegosaurus and Ankylosaurus. However, compared to some of its later relatives, Scutellosaurus was small and lightly armored. Along its back and extending onto the base of the tail were rows of small bony “shields” embedded in the skin. Some of these shields were flat, while others were pitched, like little roofs. The largest shields formed two rows that ran along the middle of the back.

Spread across the back, the shields formed a protective armored layer that would have defended the animal from attacks by such meat-eating dinosaurs as Dilophosaurus, with which it shared its early Jurassic world. Scutellosaurus would also have been reasonably fleet of foot and therefore able to escape predators by weaving its way through tangled undergrowth.

Skeletal reconstruction showing known material. Author: Jaime A. Headden

Scutellosaurus was lightly built, and was probably capable of walking on its hind legs. It had an unusually long tail, possibly to provide a counterbalance against the weight of the armored body. It was around 1.2 metres (3.9 ft) long, 50 centimetres (20 in) tall at the hips, and weighed 10 kilograms (22 lb). The fossil evidence includes several partial skeletons recovered from Arizona by the Museum of Northern Arizona and the University of California Museum of Paleontology, although the skull is poorly known from these specimens. There were several hundred osteoderms running along its neck to its back and as far down as its tail. These formed parallel rows, with as many as five rows on each side. It also had double rows of osteoderms, or external plates, running neck to tail. Some of these shields were flat, while others were pitted.

Pachyrhinosaurus

Saturday, December 10, 2016

P. perotorum mounted at the Perot Museum

Pachyrhinosaurus (meaning “thick-nosed lizard”) is an extinct genus of centrosaurine ceratopsid dinosaurs from the Late Cretaceous period of North America. The first examples were discovered by Charles M. Sternberg in Alberta, Canada, in 1946, and named in 1950. Over a dozen partial skulls and a large assortment of other fossils from various species have been found in Alberta and Alaska. A great number were not available for study until the 1980s, resulting in a relatively recent increase of interest in the Pachyrhinosaurus. Three species have been identified. P. lakustai, from the Wapiti Formation, the bonebed horizon of which is roughly equivalent age to the upper Bearpaw and lower Horseshoe Canyon Formations, is known to have existed from about 73.5-72.5 million years ago. P. canadensis is younger, known only from the lower Horseshoe Canyon Formation, about 71.5-71 Ma ago. Fossils of the youngest species, P. perotorum, have been recovered from the Prince Creek Formation of Alaska, and date to 70-69 million years ago. The presence of three known species makes this genus the most speciose among the centrosaurines.

Pachyrhinosaurus by PrehistoricWildlife.com

QUICK PACHYRHINOSAURUS FACTS

  • May have run as fast as 20 MPH

  • Lived in parts of Canada and Alaska

  • Its name means “thick nosed lizard”

  • 3 species of this dinosaur have been discovered

  • Weighed as much as a Black Rhinoceros

ABOUT PACHYRHINOSAURUS

Pachyrhinosaurus is a dinosaur which lived around 70 million years ago during the late Cretaceous Period. It was first discovered in the late 1940s by Charles M. Sternberg, son of famous paleontologists Charles H. Sternberg who found specimens in Alberta, Canada and Alaska, U.S. He also named the fossil, calling it Pachyrhinosaurus—a name which means “thick nosed lizard.”

Pachyrhinosaurus was an herbivore that was approximately 20 to 26 feet long, about 6 feet high and weighed around 2 to 3 tons. Like other dinosaur’s in this dinosaur order, the Pachyrhinosaurus had a large bony frill that came out of the back of its skull. However, what makes this dinosaur different from the other ones in it dinosaur classification is that instead of horns on its nose it had a large bony bump called a “boss.” It did have a pair of horns that grew out of the top of its frill and it may have had horns over its eyes.

Like other herbivores of this time, the Pachyrhinosaurus probably traveled in herds that kept it safe from predators. Paleontologists believe that some of these herds may have had hundreds or thousands of animals in it at one time. If this was indeed the case, then it would have made it quite difficult on predators. It is also likely that these dinosaurs also nested their eggs like modern birds and may have even taken care of them. This has been found to be the case in other dinosaur types such as the Protoceratops and the Styracosaurus.

Paleontologists believe this animal probably lived off of a diet that consisted mainly of palms and cycads and other tough plant material that it could tear off and crush with its beak. It also had cheek teeth that it could use to further masticate this tough plant material, making it easier to digest.

P. perotorum engaged in intraspecific combat. Artistic rendering of Pachyrhinosaurus perotorum engaged in head-butting/pushing behavior. In the first description of Pachryhinosaurus by Sternberg, he speculated that the enlarged nasal boss in the taxon might have been used in head battering or pushing behavior, an idea emphasized by this image of two Pachyrhinosaurus perotorum sparring with their craniofacial bosses, while a third looks on. Author: Karen Carr

Anchiceratops

Saturday, December 10, 2016

Anchiceratops by atrox1 on DeviantArt

Anchiceratops is an genus of chasmosaurine ceratopsid dinosaur that lived approximately 72 to 71 million years ago during the latter part of the Cretaceous Period in what is now Alberta, Canada. Anchiceratops was a medium-sized, heavily built, ground-dwelling, quadrupedal herbivore that could grow up to an estimated 5 m (16.4 ft) long. Its skull featured two long brow horns and a short horn on the nose. The skull frill was elongated and rectangular, its edges adorned by coarse triangular projections. About a dozen skulls of the genus have been found.

NMC 8547 mounted at the Canadian Museum of Nature, completed with a skull cast of NMC 8535. NMC 8547 might represent a separate taxon.

Anchiceratops was a medium-sized ceratopsid. If specimen NMC 8547 is not taken into account, no very exact estimations of the body length of Anchiceratops can be given. Some popular science book state that it approached 20 feet (6 m) in length. In 2010 Gregory S. Paul, on the assumption that specimen NMC 8547 represented Anchiceratops, estimated its length at 4.3 metres, its weight at 1.2 tonnes.

The first remains of Anchiceratops were discovered along the Red Deer River in the Canadian province of Alberta in 1912 by an expedition led by Barnum Brown. The holotype, specimen AMNH 5251, is the back half of a skull, including the long frill, and two other partial skulls, specimens AMNH 5259 (the paratype) and AMNH 5273, were found at the same time, which are now stored in the American Museum of Natural History in New York City. A complete skull designated NMC 8535, was discovered by Charles M. Sternberg at Morrin in 1924, and was described as A. longirostris five years later, in 1929. but this species is widely considered a junior synonym of A. ornatus today. In total, at least ten incomplete skulls have been recovered. The skulls are different with respect to their proportions (e.g. size of the supraorbital horn cores, the dimensions of the frill) which had led researchers to conclude that the disparity is a result of interspecific differences or due to sexual dimorphism.

ROM 802, a skull in the Royal Ontario Museum by Daderot

The most distinctive features of this dinosaur are in its unusual neck frill. The frill is moderately long and rectangular with small, oval fenestrae (openings). The edge of the frill is thick just behind the brow horns. On the back of the frill are six large epoccipitals (bony knobs around the frill) that were expanded into short, triangular, backward-pointing spikes. Also on the frill are two short spikes that curve up and out.

Anchiceratops had a short nasal horn, a very long nose, and two moderate-size brow horns. Its skeleton shows it had a very short tail, but otherwise it looked much like other ceratopsids.

Anchiceratops lived at the same time as its close relative Arrhinoceratops. It was also closely related to TorosaurusChasmosaurusPentaceratops, and Triceratops.

Styracosaurus

Saturday, December 10, 2016

Styracosaurus by atrox1 on DeviantArt

Styracosaurus (meaning “spiked lizard” from the Ancient Greek styrax/στύραξ “spike at the butt-end of a spear-shaft” and sauros/σαῦρος “lizard”) was a genus of herbivorous ceratopsian dinosaur from the Cretaceous Period (Campanian stage), about 75.5 to 75 million years ago. It had four to six long horns extending from its neck frill, a smaller horn on each of its cheeks, and a single horn protruding from its nose, which may have been up to 60 centimetres (2 ft) long and 15 centimetres (6 in) wide. The function or functions of the horns and frills have been debated for many years.

Size compared to a human by Marmelad

Styracosaurus was a relatively large dinosaur, reaching lengths of 5.5 metres (18 ft) and weighing nearly 3 tonnes. It stood about 1.8 meters (6 ft) tall. Styracosaurus possessed four short legs and a bulky body. Its tail was rather short. The skull had a beak and shearing cheek teeth arranged in continuous dental batteries, suggesting that the animal sliced up plants. Like other ceratopsians, this dinosaur may have been a herd animal, traveling in large groups, as suggested by bonebeds.

Named by Lawrence Lambe in 1913, Styracosaurus is a member of the Centrosaurinae. One species, S. albertensis, is currently assigned to Styracosaurus. Other species once assigned to the genus have since been reassigned elsewhere.

Styracosaurus skeleton at Canadian Museum of Nature, Ottawa, Ontario. The Vagaceratops irvinensis type specimen can be seen in the lower left corner. Author: D. Gordon E. Robertson

The first fossil remains of Styracosaurus were collected in Alberta, Canada by C.M. Sternberg (from an area now known as Dinosaur Provincial Park, in a formation now called the Dinosaur Park Formation) and named by Lawrence Lambe in 1913. This quarry was revisited in 1935 by a Royal Ontario Museum crew who found the missing lower jaws and most of the skeleton. These fossils indicate that S. albertensis was around 5.5 to 5.8 meters in length and stood about 1.65 meters high at the hips. An unusual feature of this first skull is that the smallest frill spike on the left side is partially overlapped at its base by the next spike. It appears that the frill suffered a break at this point in life and was shortened by about 6 centimeters (2 in). The normal shape of this area is unknown because the corresponding area of the right side of the frill was not recovered.

Styracosaurus is a member of the Centrosaurinae. Other members of the clade include Centrosaurus (from which the group takes its name), PachyrhinosaurusAvaceratopsEiniosaurusAlbertaceratopsAchelousaurusBrachyceratops, and Monoclonius, although these last two are dubious. Because of the variation between species and even individual specimens of centrosaurines, there has been much debate over which genera and species are valid, particularly whether Centrosaurusand/or Monoclonius are valid genera, undiagnosable, or possibly members of the opposite sex. In 1996, Peter Dodson found enough variation between CentrosaurusStyracosaurus, and Monoclonius to warrant separate genera, and that Styracosaurusresembled Centrosaurus more closely than either resembled Monoclonius. Dodson also believed one species of MonocloniusM. nasicornis, may actually have been a female Styracosaurus. However, most other researchers have not accepted Monoclonius nasicornis as a female Styracosaurus, instead regarding it as a synonym of Centrosaurus apertus. While sexual dimorphism has been proposed for an earlier ceratopsian, Protoceratops, there is no firm evidence for sexual dimorphism in any ceratopsid.

Depiction of dietary niche partitioning among megaherbivorous dinosaurs from the DPF (MAZ-2).  Left to right: Chasmosaurus belli, Lambeosaurus lambei, Styracosaurus albertensis, Euoplocephalus tutus, Prosaurolophus maximus, Panoplosaurus mirus. A herd of S. albertensis looms in the background. Image courtesy of J.T. Csotonyi.

Styracosaurus is known from the Dinosaur Park Formation, and was a member of a diverse and well-documented fauna of prehistoric animals that included horned relatives such as Centrosaurus and Chasmosaurus, duckbills such as ProsaurolophusLambeosaurusGryposaurusCorythosaurus, and Parasaurolophustyrannosaurids GorgosaurusDaspletosaurus, and armored Edmontonia and Euoplocephalus.

Tenontosaurus

Saturday, December 10, 2016

Reconstructed skeleton of Tenontosaurus, from the Cloverly Formation.

Tenontosaurus (meaning “sinew lizard”) is a genus of medium- to large-sized ornithopod dinosaur. The genus is known from the late Aptian to Albian ages of the middle Cretaceous period sediments of western North America, dating between 115 and 108 million years ago.

The genus contains two species, Tenontosaurus tilletti (described by John Ostrom in 1970) and Tenontosaurus dossi (described by Winkler, Murray, and Jacobs in 1997). Many specimens of T. tilletti have been collected from several geological formations throughout western North America. T. dossi is known from only a handful of specimens collected from the Twin Mountains Formation of Parker County, Texas.

Restoration of T. tilletti by Nobu Tamura

It was about 6.5 to 8 metres (21 to 26 ft) long and 3 metres (9.8 ft) high in a bipedal stance, with a mass of somewhere between 1 to 2 tonnes (1 to 2 short tons). It had an unusually long, broad tail, which like its back was stiffened with a network of bony tendons.

Tenontosaurus fed on plant matter, which it broke off with its horny beak and then chewed with its teeth, all of which were at the back of its mouth. The small, swift-moving Deinonychus was Tenontosaurus’s main enemy. The teeth of this predator have been found along with Tenontosaurus skeletons. Some scientists therefore believe that Deinonychus hunted the larger animal in packs, but there is no real evidence to support this. In spite of what would seem to be the advantages of having clawed feet and a huge tail, Tenontosaurus would have been easy catch for packs of sharp-fanged Deinonychus.

T. tilletti with juveniles, in front of Deinonychus, Academy of Natural Sciences, Philadelphia by Daderot

The first Tenontosaurus fossil was found in Big Horn County, Montana by an American Museum of Natural History (AMNH) expedition in 1903. Subsequent digs in the same area during the 1930s unearthed 18 more specimens, and four specimens were found during the 1940s. Despite the large number of fossil specimens, the animal was not named or scientifically described during this time, though Barnum Brown of the AMNH gave it the informal name “Tenantosaurus”, “sinew lizard”, in reference to the extensive system of stiffening tendons in its back and tail.

Torosaurus

Saturday, December 10, 2016

Torosaurus by atrox1 on DeviantArt

Torosaurus is a genus of herbivorous ceratopsid dinosaur that lived during the late Maastrichtian stage of the Cretaceous period, between 68 and 66 million years ago. Its name means “perforated lizard” in reference to the large openings in its frill. Fossils have been discovered across the Western Interior of North America, from Saskatchewan to southern Texas.

Torosaurus by PrehistoricWildlife.com

Torosaurus possessed one of the largest skulls of any known land animal. The frilled skull reached up to 2.77 metres (9.1 ft) in length. From head to tail, Torosaurus is thought to have measured about 7.6 to 9 m (24.9 to 29.5 ft) long and weighed four to six tonnes. Torosaurus is distinguished from the contemporary Triceratops by an elongate frill with large openings (fenestrae), long squamosal bones of the frill with a trough on their upper surface, and the presence of five or more pairs of hornlets (epoccipitals) on the back of the frill. Torosaurus also lacked the long nose horn seen in Triceratops prorsus, and instead resembled the earlier and more basal Triceratops horridus in having a short nose horn. Three species have been named, Torosaurus latusT. gladius and T. utahensisT. gladius is no longer considered a valid species, however.

Recently the validity of Torosaurus has been disputed. A 2010 study of fossil bone histology combined with an investigation of frill shape concluded that Torosaurus probably represented the mature form of Triceratops, with the bones of typical Triceratops specimens still immature and showing signs of a first development of distinct Torosaurus frill holes. During maturation, the skull frill would have been greatly lengthened and holes would have appeared in it. In 2011, 2012 and 2013, however, studies of external features of known specimens have claimed that morphological differences between the two genera preclude their synonymy. The main problems are a lack of good transitional forms, the apparent existence of authentic Torosaurus subadults, different skull proportions independent of maturation and the assertion that hole formation at an adult stage is not part of a normal ceratopsian maturation sequence.

Scanella & Horner saw Nedoceratops as an ontogenetic transitional form between Triceratops and Torosaurus

All ceratopsians, including Torosaurus and Triceratops, were herbivores. The jaws contained rows of teeth that slid past each other as the jaws closed, creating a shearing action that would have allowed Torosaurus to process tough vegetation, perhaps the ferns that formed much of the ground cover. A slightly longer, narrower snout in Torosaurus than in Triceratops suggests that they might have had different feeding preferences.

Tuojiangosaurus

Saturday, December 10, 2016

Tuojiangosaurus

Tuojiangosaurus (meaning “Tuo River lizard”) is a genus of herbivorous stegosaurid dinosaur from the Late Jurassic Period, recovered from the Upper Shaximiao Formation of what is now Sichuan Province in China.

Tuojiangosaurus has the typical narrow and low head, bulky body, and low teeth of other stegosaurids. The limbs, especially the arms, are rather short. There are at least twenty-five dentary teeth. The teeth have a thick base, cingulum, merging at the inside into a triangular vertical median ridge. The dorsal vertebrae have tall neural arches. The shoulderblade has a rectangular acromion.

Tuojiangosaurus multispinus by Pachyornis

Tuojiangosaurus was by Dong placed in Stegosauridae in 1977, more precisely in the Stegosaurinae. In 2004, a cladistic analysis by Galton recovered Tuojiangosaurus in a rather derived position, as a sister species of Chialingosaurus. An analysis by Susannah Maidment in 2006, confirmed its status as a member of the Stegosauridae.

Like KentrosaurusTuojiangosaurus had two rows of plates along the spine, which became taller over the hip region. Those at the neck and front trunk were rounded or pear-shaped; the plates at the rear back became more triangular and pointed. All plates have a thickened central section, as if they were modified spikes. Dong estimated there were about seventeen pairs of plates and spikes. Tuojiangosaurus had at least two outward-pointing, rather robust, spikes on each side of the end of the tail, angled at approximately 45 degrees to the vertical. In stegosaurids, this spike arrangement has become affectionately known as the “thagomizer”. Dong thought it were possible that there were four pairs of spikes. Paul, based on “Chungkingosaurus sp. 3″ specimen CV 00208, interpreted the thagomizer as a “pin-cushion array”, with two vertical pairs of thick spikes and a third pair of narrow spikes pointing to behind.

Reconstructed skeleton of Tuojiangosaurus

Another two stegosaurs-Chialingosaurus and Chungkingosaurus-lived in China alongside Tuojiangosaurus in the late Jurassic. Both of these were smaller than Tuojiangosaurus, and they may have coexisted by dividing up the plants they ate based on the height at which they fed. Alternatively, the smaller stegosaurs may have inhabited more enclosed forests, which would have protected them from predators, while the larger Tuojiangosaurus lived in a more open environment, protected by its size.

In 1974, during construction of the Wujiaba dam in Zigong, Sichuan, the remains of stegosaurians were found.

The type and only species of TuojiangosaurusTuojiangosaurus multispinus, was named and described in 1977 (exactly a hundred years after the naming of Stegosaurus by Othniel Charles Marsh) by Dong Zhiming, Zhou Shiwu, Li Xuanmin and Chang Yijong. The generic name is derived from the River (jiang) Tuo. The specific name is derived from Latin multus, “many”, and spina, “spine”.

Yangchuanosaurus attacking Tuojiangosaurus. Author: Foolp

The holotype, CV 209, was found in a layer of the Upper Shaximiao Formation, dating from the Oxfordian-Kimmeridgian. It consists of a rather complete skeleton that however lacks parts of the skull., lower jaws, tail and limbs. In 1977, it represented the most complete stegosaurian skeleton found in Asia. The paratype was specimen CV 210, a sacrum. Subsequently, more material has been referred, including juveniles. This complemented the holotype with elements of the skull, especially the braincase, and the lower jaws.

A mounted skeleton of Tuojiangosaurus multispinus is on display at the Municipal Museum of Chongqing. In addition, a mounted cast is on display at the Natural History Museum, in London.

Troodon

Saturday, December 10, 2016

Troodon by Taena Doman

Troodon is a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Cretaceous period (about 77 mya). It includes at least one species, Troodon formosus, known from Montana. Discovered in October 1855, T. formosus was among the first dinosaurs found in North America, although it was thought to be a lizard until 1877. Several well-known troodontid specimens from the Dinosaur Park Formation in Alberta were once believed to be members of this genus. However, recent analyses in 2017 have found the genus to be undiagnostic and referred some of these specimens to the genus Stenonychosaurus (long believed to be synonymous with Troodon) and others to the newly created genus Latenivenatrix.

Troodon by Prehistoric-Wildlife.com

The genus name is Greek for "wounding tooth", referring to the teeth, which were different from those of most other theropods known at the time of their discovery. The teeth bear prominent, apically oriented serrations. These "wounding" serrations, however, are morphometrically more similar to those of herbivorous reptiles, and suggest a possibly omnivorous diet.

The name was originally spelled Troödon (with a diaeresis) by Joseph Leidy in 1856, which was officially amended to its current status by Sauvage in 1876. The type specimen of Troodon has caused problems with classification, as the entire genus is based only on a single tooth from the Judith River Formation. Troodon has historically been a highly unstable classification and has been the subject of numerous conflicting synonymies with similar theropod specimens.

The Troodon tooth was originally classified as a "lacertilian" (lizard) by Leidy, but reassigned as a megalosaurid dinosaur by Nopcsa in 1901 (Megalosauridae having historically been a wastebin taxon for most carnivorous dinosaurs). In 1924, Gilmore suggested that the tooth belonged to the herbivorous pachycephalosaur Stegoceras, and that Stegoceras was in fact a junior synonym of Troodon (the similarity of troodontid teeth to those of herbivorous dinosaurs continues to lead many paleontologists to believe that these animals were omnivores). The classification of Troodon as a pachycephalosaur was followed for many years, during which the family Pachycephalosauridae was known as Troodontidae. In 1945, Charles Mortram Sternberg rejected the possibility that Troodon was a pachycephalosaur due to its stronger similarity to the teeth of other carnivorous dinosaurs. With Troodon now classified as a theropod, the family Troodontidae could no longer be used for the dome-headed dinosaurs, so Sternberg named a new family for them, Pachycephalosauridae.

The first specimens assigned to Troodon that were not teeth were both found by Sternberg in the early 1930s, in the Dinosaur Park Formation of Alberta. The first was named Stenonychosaurus inequalis by Sternberg in 1932, based on a foot, fragments of a hand, and some tail vertebrae. A remarkable feature of these remains was the enlarged claw on the second toe, which is now recognized as characteristic of early paravians. Sternberg initially classified Stenonychosaurus as a member of the family Coeluridae. The second, a partial lower jaw bone, was described by Gilmore (1932) as a new species of lizard which he named Polyodontosaurus grandis. Later, in 1951, Sternberg recognized P. grandis as a possible synonym of Troodon, and speculated that since Stenonychosaurus had a "very peculiar pes" and Troodon "equally unusual teeth", they may be closely related. Unfortunately, no comparable specimens were available at that time to test the idea. In a recent revision of the material by van der Reest & Currie, Polyodontosaurus was determined to be a nomen dubium, not fit fot synonymy with other taxa.

Skeletal restoration of T. inequalis by Scott Hartman

Troodon is considered to be one of the most derived members of its family. Along with Zanabazar, Saurornithoides and Talos it forms a clade of specialized troodontids.

One study was based on multiple Troodon teeth that have been collected from Late Cretaceous deposits from northern Alaska. These teeth are much larger than those collected from more southern sites, providing evidence that northern Alaskan populations of Troodon grew to larger average body size. This study also provides an analysis of the proportions and wear patterns of a large sample of Troodon teeth. It proposes that the wear patterns of all Troodon teeth suggest a diet of soft foods - inconsistent with bone chewing, invertebrate exoskeletons, or tough plant items. This study hypothesizes a diet primarily consisting of meat.

Troodon Skull

The type specimen of Troodon formosus was found in the Judith River Formation of Montana. The rocks of the Judith River Formation are equivalent in age with the Oldman Formation of Alberta, which has been dated to between 77.5 and 76.5 million years ago.

Source: www.wikipedia.org / www.natgeo.com

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